Here is a spreadsheet to determine what level will provide the most profits for releasing wolves! This assumes that you don't care about knucklebone tokens and you only want pebbles:
https://docs.google.com/spreadsheets/d/1I8dcQzDMPBw76pUIS6iLm-1wusBLOfZNhdXafLNvTu4/edit#gid=0
Preview of the sheet, with wolf prices at 2kp:
To find the most efficient level, look in the total pebbles per exp column and find the highest number. Then see what level that is. I may add a box that does this for you at some point but I'm not sure what formula to use and I have some wolves to go train to level 6